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[¥H¥Øªº½×½Íºt¤Æ¨ì©³¦³¤°»ò°ÝÃD¡H]Is?~E ©©øÂν׾ -- ©øÂν׾¡@¡@ nV ¬°¤F¡u¾AªÌ¥Í¦s¡v¡A¥Íª«¦ÛµMµo®i¥X³\¦hÁ׼ĵ¦²¤¡A¬°¤F¦Û½Ã¡A§ó¬°¤Fª«ºØªº©µÄò¡C(¸`¦Û¥þ°ê¤½¨p¥ß°ª¯Å¤¤¾Ç106¾Ç¦~«×¾Ç¬ì¯à¤O´úÅç²Ä¤T¦¸Áp¦X¼ÒÀÀ¦Ò¸Õ¤§°ê¤å¦Ò¬ì«D¿ï¾ÜÃD²Ä¤@ÃD°Ñ¦Ò½d¤å)qqx&7h ©©øÂν׾ -- ©øÂν׾¡@¡@ | ¾Ç´ú°ê¤å¼g§@ÃD¥Ø»P¦ÛµM¬ìµ²¦XÅkµM¤w¦¨ÁͶաA¦ý³o¼Ëªº¸ó¾Ç¬ì¾ã¦X¶Õ¥²·|¹J¨ì§xÃø¡A¤°»ò§xÃø¡H¥XÃD¦Ñ®v¦³¨S¦³·d²M·¡¤£¦P»â°ì¶¡ªºª¾ÃѸ¨®t¡C²Ä¤T¦¸¥þ°ê°ª¤¤Áp¦X¼ÒÀÀ¦Ò°ê¤å¦Ò¬ì«D¿ï¾ÜÃD²Ä¤@ÃD¡AÃD·F¬D¤F¤T½g»P¥Íª«Á׼Ħ欰¬ÛÃöªº¿ï¤å¡A¨Ã½Ð¦Ò¥Í¥H¡u¥Íª«Á׼Ħ欰µ¹§Úªº±Òµo¡v¬°ÃD¼¶¼g¤@½g¤å³¹¡C¦Ò¸Õµ²§ô«á°Ñ¦Ò½d¤å¥X¨Ó¡A½d¤å¤@¶}©l´N¥H¡G¡y¬°¤F¡u¾AªÌ¥Í¦s¡v¡z¶}ÀY¡A³o¦³¤°»ò°ÝÃD¡H³o´N¬O«D±`¨å«¬ªº¥H¥Øªº½×½Íºt¤Æ¡A½Ð¦U¦ì¸Õ·Qºt¤Æ¬O«ç»òµo¥Íªº¡H¥Íª«¥i¥H¦³·NÃѪº¨M©w·Qnºt¤Æ¦¨¤°»ò¼Ë¤l¶Ü¡HÁÙ¬O¤Ñ¾Üªº¤O¶q¨Ï±o¥Íª«ºt¤Æ¦¨²{¦b³oӼˤl¡HgOD ©©øÂν׾ -- ©øÂν׾¡@¡@ >Ry.!k §ÚÌ¥H¥É±a»ñ½º(Papilio polytes)¤Î¬õ¯]»ñ½º(Pachliopta aristolochiae)¤§¶¡ªºÀÀºAÃö«Y¬°¨Ò¡A¥É±a»ñ½º¬O«D±`¸g¨åªº¨©¤óÀÀºAªº®×¨Ò¡A¨e¬O©úÅ㪺»Û©Ê¦h«¬©Ê(female polymorphism)ª«ºØ¡A¦b»OÆW¥iµo²{»ÛÂΦ³¨âºØ«¬ºA¡A¤@ºØ¬OÀÀºA«¬(form polytes)¥t¤@ºØ¬°«DÀÀºA«¬(form cyrus)¡AÀÀºA«¬ÀÀºAªºmodel¬°¬õ¯]»ñ½º¡A±°¹ªÌ¥i¯à¦]¬°¦Y¹L¤£¦n¦Yªº¬õ¯]»ñ½º¦Ó°§CÀÀºA«¬¥É±a»ñ½ºªº³Q®·¹¾÷²v¡A¦Ó«DÀÀºA«¬«h¦h³Q»{¬°¯à°÷§l¤Þ¶¯½º¡C1 ¸ÕµÛ¥H¥Øªº½×¤ÎÀÀ¤H¤Æªº¤f§k¨Ó½Í³oÓijÃD·|ÅÜ«ç¼Ë¡Hdy8} ©©øÂν׾ -- ©øÂν׾¡@¡@ ¥É±a»ñ½º¬°¤F¬¡¤U¥h¡A¯ª©v¤Q¤K¥N³£¶}©l¾ÇµÛ³ì¸Ë¦¨¦³¬rªº¬õ¯]»ñ½º¡A¨e̬°¤F°kÁ×±°¹ªÌªº°lÀ»¡A©Ò¥H½º½º³£¥²¶·¾Ç²ß¬õ¯]»ñ½ºªº¼Ë¤l¡A¾Ç±o¤£¹³±o´N¬O¨e¬¡¸Ó¡Aµ¥µÛ³Q±°¹ªÌ¦Y±¼¡A¤@¥N¹L¤F¤@¥N¡AÀÀºA«¬´N¥X²{¤F¡C±q³o¼Ë¤U¥h½Í¤£§K¤S·|¶]¦^©Ô°¨§Jªº¥Î¶i¼o°h»¡²z½×¡CN]T7^ ©©øÂν׾ -- ©øÂν׾¡@¡@ DvC,_d ¦ý¹ê»Ú¤W¥É±a»ñ½º¯Í¤Wªº´³¯¾¬O¥Ñ¤@Ó¥s°µdoublesexªº¶W°ò¦](supergene)©Ò±±¨î¡C¦¦b50-60¦~«eClarke©MSheppard´Nµoªí¦h½g³ø§i(Clarke and Sheppard,1960; Clarke et al., 1968; Clarke and Sheppard, 1972)¡A³z¹LÂø¥æ¹êÅç¡A´£¥X¥É±a»ñ½ºµ¥»ñ½º¯Í¤Wªº´³¯¾½Õ±±°ò¦]¬OÓ¶W°ò¦]¡A¦Ü2014¦~¬ì¾Ç®a¤~§ä¨ì³o¬O¥Ñ¤@Ó¥s°µdoublesexªº¶W°ò¦](Krushnamegh et al., 2014)¡A¦Ó2015¦~«h¤S¦³¤@½g³ø§i«ü¥X¥É±a»ñ½ºÀÀºA«¬ªº¥X²{¬Odoublesexªº°ò¦]§Ç¦CËÂà©Ò¾ÉP(Hideki et al., 2015)¡CMAn ©©øÂν׾ -- ©øÂν׾¡@¡@ 971] ©Ò¥H»¡¡Aºt¤Æ¬O¦³¥Øªºªº¶Ü¡H½Ö¯à¦³·NÃѪº¾Þ±±¦Û¤vªº°ò¦]¥s¥¦¤£n½Õ±±¦Û¤vªº©Êª¬©O¡H9|] ©©øÂν׾ -- ©øÂν׾¡@¡@ -`L~ ©©øÂν׾ -- ©øÂν׾¡@¡@ ? ¦¹¥DÃD¬ÛÃö¹Ï¤ù¦p¤U¡G; 94S qS «ö¦¹¬d¬Ý¹Ï¤ù¸Ô²Ó¸ê®Æ¥¿¦bŪ¨ú¦¹¹Ï¤ùªº¸Ô²Ó¸ê®Æ¡A½ÐµyÔ ... | }q$ ©©øÂν׾ -- ©øÂν׾¡@¡@ +RB" ©©øÂν׾ -- ©øÂν׾¡@¡@ ! ¥É±a»ñ½ºÀÀºA«¬»ÛÂÎU ©©øÂν׾ -- ©øÂν׾¡@¡@ k=5& °Ñ¦Ò¤åÄm¡GLP>W C. A. Clarke, Philip Macdonald Sheppard (1960) Super-gene and mimicry. Heredity, 14: 175-185.p ©©øÂν׾ -- ©øÂν׾¡@¡@ R!'^ C. A. Clarke, Philip Macdonald Sheppard, I. W. B. Thornton (1968) The genetics of the mimetic butterfly Papilio memnon L. Philosophical Transactions of the Royal Society B, 254: 37-89.i ©©øÂν׾ -- ©øÂν׾¡@¡@ 0e C. A. Clarke, Philip Macdonald Sheppard (1972) The genetics of the mimetic butterfly Papilio polytes L. Philosophical Transactions of the Royal Society B, 263: 431-458.\_=4nY ©©øÂν׾ -- ©øÂν׾¡@¡@ +Ag) Hideki Nishikawa, Takuro Iijima, Rei Kajitani, Junichi Yamaguchi, Toshiya Ando, Yutaka Suzuki, Sumio Sugano, Asao Fujiyama, Shunichi Kosugi, Hideki Hirakawa, Satoshi Tabata, Katsuhisa Ozaki, Hiroya Morimoto, Kunio Ihara, Madoka Obara, Hiroshi Hori, Takehiko Itoh, Haruhiko Fujiwara (2015) A genetic mechanism for female-limited Batesian mimicry in Papilio butterfly. Nature genetics, 47: 405-409.\z$a ©©øÂν׾ -- ©øÂν׾¡@¡@ e Krushnamegh Kunte, Wei Zhang, Robert D. Reed, Sean P Mullen (2014) Doublesex is a mimicry supergene. Nature, 507: 229-232.u4'@O ©©øÂν׾ -- ©øÂν׾¡@¡@ e Toshio Sekimura, Yuta Fujihashi, Yasuhiro Takeuchi (2014) A model for population dynamics of the mimetic butterfly Papilio polytes in Sakishima Islands, Japan. Journal of theoretical biology, 361: 133-140.J
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